Plot: Remy, a provincial rat with a wonderful sense of smell, hates garbage and risks death to enter a human kitchen where he discovers real food and the cooking of five-star chef, Anton Gusteau, author of "Anyone Can Cook". On the day Remy learns his hero has died, he is evicted and ends up alone in Paris. By luck, he discovers Gasteau's restaurant, down to three stars and run by a frozen-food-hawking chef. As Remy enters, so does Linguini, a clumsy youth hired as a garbage boy. To save the soup that Linguini accidentally fouls, Remy throws in some ingredients; the soup is a success and Linguini's career as a chef is born. Can Remy find a way to maintain the fiction and use his gift?
Alternative Plot: Remy (Patton Oswalt), a resident of Paris, appreciates good food and has quite a sophisticated palate. He would love to become a chef so he can create and enjoy culinary masterpieces to his heart's delight. The only problem is, Remy is a rat. When he winds up in the sewer beneath one of Paris' finest restaurants, the rodent gourmet finds himself ideally placed to realize his dream.
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